GIES Case Study on Fusong Lushuihe Korean Pine Seeds Forest Ecosystem

QI Wei1,2  ZHU Jianhua3  WANG Gang4  GAO Jiantang5  DU Yuchuan5  ZHAO Wenxu6 WANG Xiuqin7  LIU Huiqiang7  JIN Xin7  XU Yanguo8  ZHANG Zenggui9  CHEN Yongting3 LIU Yan3  LIU Yubo3  NIU Ziming3  ZHU Wenhao1,2  LI Yu1,2  FAN Kai1,2  TANG Xinrui1,2

1 Institute of Geographic Sciences and Natural Resources Research, Chinese Academy of Sciences, Beijing 100101, China

2 University of Chinese Academy of Sciences, Beijing 100049, China

3 Northeast Normal University, Changchun 130000, China

4 People's Government of Fusong County, Baishan City, Jilin Province, Fusong County 134500, China

5 Fusong County Market Supervision and Administration Bureau, Baishan City, Jilin Province, Fusong County 134500, China

6 Lazihe Village, Lushui Town, Fusong County, Baishan City, Jilin Province, Fusong County 134500, China

7 Jilin Changbai Mingzhu Forest Food Co., Ltd., Fusong County 134500, China

8 Hongguang Forest Farm of Lushuihe Forestry Co., Ltd., Fusong County 134500, China

9 Fusong County Lushuihe Hualong Chinese Herbal Medicine Planting Professional Cooperative, Fusong County 134500, China

 

Abstract: The core production area of Lushuihe Korean Pine Seeds is located in Lushuihe Town, Fusong County, Jilin Province, within the central region of the Changbai Mountains. The Lushuihe Pinus koraiensis forest area, endowed with unique natural environmental conditions such as mid-to-high altitude, high vegetation coverage, cold-temperate humid climate, and fertile volcanic ash soil, endows the pine nut kernels with plumpness, milky-white color, and rich aroma, making them a natural food of high nutritional value. This study analyzes Lushuihe Town, the case area, from the perspectives of the ecological environment of the Pinus koraiensis forest, product characteristics, management features, and socioeconomic development, and summarizes the habitat protection and sustainable development model of the Lushuihe Korean Pine Seed production area. The case dataset mainly consists of data on the spatial scope of the case area, physical geographical conditions, product characteristics of Lushuihe Korean Pine Seed kernels, socioeconomic development, management and culture, etc. The dataset is stored in formats such as .shp, .tif, .xlsx, .docx, and .jpg, consisting of 82 data files with a data volume of 264 MB (compressed into one file of 77.3 MB).

Keywords: Fusong; Lushuihe; Korean pine seeds; Pinus koraiensis forest; GIES; Case 36

DOI: https://doi.org/10.3974/geodp.2026.03.08

CSTR: https://cstr.escience.org.cn/CSTR:20146.14.2026.03.08

1 Introduction

Pinus koraiensis, also known as the “fruit pine,” is a tall arborescent species belonging to the division Gymnospermae, family Pinaceae, genus Pinus, subgenus Strobus. It is an edifier species of the coniferous and broad-leaved mixed forest, which is one of the zonal vegetation types in the temperate zone of Northeast China. The species is primarily distributed in the Changbai Mountains, Wanda Mountains, Zhangguangcai Range, and Xiaoxing’an Mountains in Northeast China. The natural maternal forest of P. koraiensis in Lushuihe Town, Fusong County, Jilin Province, represents the largest, most intact, and most genetically diverse natural maternal forest of P. koraiensis in China [1]. Within this maternal forest, century-old trees are common, and a specimen known as the “King of Pinus koraiensis,” with an estimated age of approximately 550 years, is also preserved. As a tall tree species, mature P. koraiensis typically reaches heights of 20–30 m, with a maximum of up to 50 m. Its crown is conical when young and gradually becomes open umbrella-shaped or tower-shaped with age. The species exhibits slow growth and a prolonged natural regeneration cycle. China’s forestry authorities have implemented the Natural Forest Protection Program (NFPP), under which wild P. koraiensis resources have been conserved and restored [2]. Wild P. koraiensis grows slowly and does not begin to bear cones until approximately 50 years of age. A single tree at full production can yield 600–800 cones annually, producing 40–60 kg of pine nuts.

The origin of Fusong Lushuihe pine nuts is Lushuihe Town, Fusong County, Jilin Province, China. The core production area is located in the hinterland of the Changbai Mountains [3], representing a premium quality pine nut in Asia [4]. Fusong Lushuihe pine nut kernels are plump, with a natural milky white color and a high kernel yield, far surpassing the quality of ordinary pine nut kernels. They are an important representative of the National Geographical Indication Products of the Changbai Mountain region. The Omega-3 and Omega-6 fatty acids in the kernels synergistically regulate cholesterol and prevent arteriosclerosis; vitamin E and anthocyanins provide dual antioxidant effects, inhibiting free radical damage; high protein and slow-release carbohydrates offer sustained satiety, making them suitable for individuals with high mental workload. In traditional Chinese medicine, they are used to moisten dryness and relieve cough, alleviate constipation, and support recovery from postpartum weakness. Fusong Lushuihe pine nuts are not only an important component of the traditional dietary culture of the Changbai Mountain region but also align with the modern concept of "natural, nutritious, and functional" healthy eating, serving as a benchmark ingredient in the high-end healthy food market [5].

The Pinus koraiensis pine nuts from Lushuihe Town, Fusong County, growing in a uniquely distinctive regional context and exclusive ecological environment, are not only a concentrated manifestation of the local natural resource endowment and historical-cultural traditions, but also a core industrial source supporting regional economic development and ensuring residents’ income. Protecting the habitat of pine nut‑producing Pinus koraiensis forests is of great significance for maintaining ecological balance, promoting local economic development, and preserving cultural traditions[6]. This study will focus on diverse information—including the natural geographical characteristics, current ecological environment status, pine nut product properties, industrial development and management models, socio‑economic development data, and historical and traditional context of this region—to conduct a systematic review and in‑depth analysis, exploring coordinated pathways between the habitat protection of Fusong Lushuihe pine nuts and sustainable development, thereby providing scientific support for the conservation practices of Pinus koraiensis forests and the high‑quality development of the pine nut industry.

2 Metadata of the Dataset

The metadata information of the "Fusong Lushuihe Korean Pine Seeds Pinus koraiensis Forest Geographical Indication Habitat Case Dataset" [7], including name, authors, geographical region, data period, data volume, data format, and dataset composition, is shown in Tab.1.

Tab.1 Metadata summary table of the "Fusong Lushuihe Korean Pine Seeds Pinus koraiensis Forest Geographical Indication Habitat Case Dataset"

Items

Description

Dataset full name

Fusong Lushuihe Korean Pine Seeds Pinus koraiensis Forest GI Habitat Case Dataset

Dataset short name

LushuiheKoreanPineSeedsCase36

Authors

QI Wei,Institute of Geographic Sciences and Resources, Chinese Academy of Sciences,qiwei@igsnrr.ac.cn

ZHU Jianhua, Northeast Normal University,zjh198827@163.com

WANG Gang,People's Government of Fusong County, Baishan City, Jilin Province,sjjbgs2024@163.com

GAO Jiantang,Fusong County Market Supervision and Administration Bureau, Baishan City, Jilin Province,sjjbgs2024@163.com

DU Yuchuan,Fusong County Market Supervision and Administration Bureau, Baishan City, Jilin Province,84809029@qq.com

ZHAO Wenxu,Lazihe Village, Lushui Town, Fusong County, Baishan City, Jilin Province,bssonglin@163.com

WANG Xiuqin,Jilin Changbai Mingzhu Forest Food Co., Ltd., bssonglin@163.com

LIU Huiqiang,Jilin Changbai Mingzhu Forest Food Co., Ltd., bssonglin@163.com

JIN Xin,Jilin Changbai Mingzhu Forest Food Co., Ltd., bssonglin@163.com

XU Yanguo,Hongguang Forest Farm of Lushuihe Forestry Co., Ltd.,bssonglin@163.com

ZHANG Zenggui,Fusong County Lushuihe Hualong Chinese Herbal Medicine Planting Professional Cooperative, bssonglin@163.com

CHEN Yongting, Northeast Normal University,3118078674@qq.com

LIU Yan, Northeast Normal University,liuyan0227@foxmail.com

LIU Yubo, Northeast Normal University,2276359635@qq.com

NIU Ziming, Northeast Normal University,13293223632@163.com

ZHU Wenhao,Institute of Geographic Sciences and Resources, Chinese Academy of Sciences,zhuwenhao18@mails.ucas.ac.cn

LI Yu,Institute of Geographic Sciences and Resources, Chinese Academy of Sciences,liyu6917@igsnrr.ac.cn

FAN kai,Institute of Geographic Sciences and Resources, Chinese Academy of Sciences,fankai0308@gmail.com

TANG Xinrui,Institute of Geographic Sciences and Resources, Chinese Academy of Sciences,tangxinrui20@mails.ucas.ac.cn

Geographical region

Lushuihe Town, Fusong County, Baishan City, Jilin Province127°25E–128°05E, 42°20N–42°45N

Year

2025

Data format

.xlsx.shp.tif.jpg.docx

Data size

246 MB (compressed into one file of 77.3 MB

Data files

spatial scope of the case area, physical geographical conditions, product characteristics of Fusong Lushuihe Korean Pine Seeds, and socioeconomic development, management and culture

Foundation

Jilin Provincial Administration for Market Regulation (2025)

Data publisher

Global Change Research Data Publishing & Repository, http://www.geodoi.ac.cn

Address

No. 11A, Datun Road, Chaoyang District, Beijing 100101, China

Data sharing policy

(1) Data are openly available and can be free downloaded via the Internet; (2) End users are encouraged to use Data subject to citation; (3) Users, who are by definition also value-added service providers, are welcome to redistribute Data subject to written permission from the GCdataPR Editorial Office and the issuance of a Data redistribution license; and (4) If Data are used to compile new datasets, the ‘ten per cent principal’ should be followed such that Data records utilized should not surpass 10% of the new dataset contents, while sources should be clearly noted in suitable places in the new dataset[8]

Communication and searchable system

DOICSTRCrossrefDCICSCDCNKISciEngineWDSGEOSSPubScholar CKRSC

3 Case Data Development

3.1 Geographical Scope Data of the Study Area

Lushuihe Town is located in the southeastern part of Jilin Province, in the northern part of Fusong County, Baishan City. It lies in the hinterland of the Changbai Mountains, within the vast forest region of the upper reaches of the Songhua River, between 127°25′E–128°05′E and 42°20′N–42°45′N (Fig.1). Lushuihe Town borders Liangjiang Town and Erdaobaihe Town of Antu County to the east, Quanyang Town to the south, Beigang Town to the west, and Yanjiang Township to the north. Lushuihe Town administers six communities: Zhanxi, Hebei, Dongshan, Beishan, Hongwei, and Xishan, as well as five administrative villages: Lazihe, Changsheng, Xinxing, Dongsheng, and Qingshuihe. In 2025, the town's administrative area is 855.58 km², with a registered population of 28,000 people.

 

Fig.1 Geographical location map of the case area

3.2 Ecological Environment Data

3.2.1 Topography and Geomorphology

The overall topography of Lushuihe Town presents a pattern of higher elevation in the south and lower elevation in the north. Based on the 90 m DEM data from SRTM[1], the authors analyzed the topographic characteristics of Lushuihe Town and found that the elevation of the town area is mainly distributed between 500–1,000 m (Fig.2). Among these, river valley lowlands with elevations below 500 m are concentrated in the northern and northwestern parts. The hilly areas at 500–700 m are the most dominant topographic type in the town. Low-mountain hilly areas at 700–900 m and locally above 900 m are mainly distributed in the central and southern parts. Flat areas with slopes <3° are distributed in strips along the river valleys. Gentle hilly slopes of 3–15° widely cover the central and eastern parts of the town. Low-to-medium mountain areas with slopes >15° are concentrated along the western margin. The moderate elevation of Lushuihe Town is consistent with the preference of Pinus koraiensis, and the extensive gentle slopes provide suitable conditions for its growth, drainage, and light selection.

Fig.2 Elevation and slope classification map of Lushuihe Town

3.2.2 Climatic Conditions

Lushuihe Town has a mid-temperate humid monsoon climate, significantly regulated by the topographic effects of the Changbai Mountains. High-temperature areas are mainly distributed in river valleys and lower-elevation zones. In the southern mountainous areas, due to higher elevation and good vegetation coverage, the surface temperature is relatively lower. The patterns of precipitation and relative humidity are generally consistent, showing higher values in the northwest and lower values in the southeast (Fig.3). Theannual average temperature of Lushuihe Town ranges from 0.9°C to 1.5°C, with maximum temperatures between 29.5°C and 32.2°C, and minimum temperatures between -39°C and -44°C. The average annual frost-free period is approximately 105 days[4]. Overall, Lushuihe Town is characterized by a cool, humid climate with sufficient accumulated temperature and large diurnal temperature variation, making it an ideal environment for the growth and nutrient accumulation of Pinus koraiensis.

a                             (b)

(c)                               (d)

(e)                                 (f)

Fig.3 Analysis of climatic conditions in Lushuihe Town (2025) (a precipitation, b temperature, c evaporation, d relative humidity, e atmospheric pressure, f windspeed)

3.2.3 Water Quality Conditions

Lushuihe Town is rich in water resources. It belongs to the upper reaches of the Songhua River water system and is one of the important tributary source and conservation areas of the Songhua River[9]. The growth of Pinus koraiensis mainly relies on natural precipitation. Therefore, this study only set up one sampling point in Dongsheng Forest Farm, where Pinus koraiensis is relatively densely distributed (Fig.4), to collect river water. The water samples were submitted to the Earthworm Soil Testing Laboratory (Shandong) Co., Ltd. for testing. The specific testing indicators included 15 items: pH, total salt content, chloride, suspended solids, chemical oxygen demand (COD), five-day biochemical oxygen demand (BOD₅), total mercury, total arsenic, cadmium, lead, hexavalent chromium, fecal coliform count, ascariasis egg count, sulfide, and anionic surfactant. The water sample test results showed that all tested items were below the concentration limits of the national irrigation water standard [10] (Tab.2).

Fig.4 Distribution map of water and soil sampling sites in the case area

Tab.2 Statistical table of surface water quality test results in the case area

Number

Pollutant item

Unit

Limit value

(≤)

Testing result

1

pH (Hydrogen ion concentration)

5.58.5

7.4

2

Total Salt Content

Mg/L

1000

65

3

Chlorides

mg/L

350

2.78

4

Suspended Solids (SS)

mg/L

100

ND

5

Chemical Oxygen Demand (COD)

mg/L

200

22

6

5-day Biochemical Oxygen Demand (BOD)

mg/L

100

4.9

7

Total Mercury (Hg)

mg/L

0.001

ND

8

Total Arsenic (As)

mg/L

0.1

ND

9

Cadmium (Cd)

mg/L

0.001

ND

10

Lead (Pb)

mg/L

0.2

ND

11

Hexavalent Chromium (Cr⁶⁺)

mg/L

0.1

ND

12

Fecal Coliform Count

MPN/L

4000

ND

13

Ascaris Egg Count

/10L

20

ND

14

Sulfides

mg/L

1

0.0427

15

Anionic Surfactants

mg/L

8

ND

 

3.2.4 Soil Conditions

The parent materials of soil formation in Lushuihe Town are mainly granite and basalt, with some sedimentary rocks. The soil types are predominantly brown forest soil and albic soil, with a general soil layer thickness of more than 0.5 m. The soils in Lushuihe Town are formed by long-term weathering and development of volcanic eruptive materials and forest litter. The surface layer is loose and porous with a thick humus layer, and volcanic sand and gravel deposits are often visible in the lower layer. The soil is generally dark brown to brownish-black in color, with high organic matter content and strong water and nutrient retention capacity, making it very suitable for the growth of Pinus koraiensis and understory characteristic crops.

3.2.4 Soil Conditions

The parent materials of soil formation in Lushuihe Town are mainly granite and basalt, with some sedimentary rocks. The soil types are predominantly brown forest soil and albic soil, with a general soil layer thickness of more than 0.5 m. The soils in Lushuihe Town are formed by long-term weathering and development of volcanic eruptive materials and forest litter. The surface layer is loose and porous with a thick humus layer, and volcanic sand and gravel deposits are often visible in the lower layer. The soil is generally dark brown to brownish-black in color, with high organic matter content and strong water and nutrient retention capacity, making it very suitable for the growth of Pinus koraiensis and understory characteristic crops.

In this study, nine soil profile sampling sites were established in the case area (Fig.4). The sampling sites were selected in forest farms with relatively dense Pinus koraiensis distribution, with full consideration given to factors such as different elevations and slope aspects. In addition, to account for the impact of human activities, soil sampling sites were also set up in Pinus koraiensis-dense areas around the town center of Lushuihe Town. Soil samples were collected in layers at depths of 020 cm, 2040 cm, 4060 cm, 6080 cm, and 80100 cm (Fig.5) to test the soil fertility and environmental conditions of the case area. The testing items included pH, organic matter, cation exchange capacity, total nitrogen, available phosphorus, readily available potassium, lead, cadmium, chromium, arsenic, mercury, copper, zinc, nickel, and organochlorine pesticide residues.

土样组合

Fig.5 Profile diagram of soil sampling sites in the case area

(1) Soil fertility status

The test results of soil profile samples showed that the pH value in this area ranged from 5.0 to 6.8, and the soil was generally acidic to weakly acidic, which falls within the suitable acidity range for the growth of Pinus koraiensis. As a core indicator for evaluating soil nutrient retention capacity, the cation exchange capacity (CEC) of the soil ranged from 8.98 to 31.84 cmol/kg, with an average value of 17.23 cmol/kg, indicating that the soil in this area has good nutrient retention capacity. Among them, the surface CEC value at sampling site P9 reached 31.84 cmol/kg, demonstrating excellent nutrient holding capacity. The CEC value showed little variation with soil depth, indicating that each soil layer possesses good nutrient retention performance. This favorable nutrient retention characteristic creates advantageous conditions for nutrient absorption by the deep root system of Pinus koraiensis, helping the tree roots to more effectively uptake nutrients from the soil, thereby promoting the healthy growth and development of Pinus koraiensis.

The soil fertility indicators at different soil layers for each sampling site are shown in Tab.3 and Fig.6. The soil organic matter content ranged from 5.29 to 89.3 g/kg, with an average value of 24.54 g/kg. The organic matter content decreased with increasing soil depth. The surface soil organic matter content was higher than that of other soil layers, showing obvious surface enrichment, while the deep soil organic matter tended to be stable but with the lowest content. The total nitrogen content varied consistently with organic matter, ranging from 0.047% to 0.498%, with an average value of 0.151%. The differences in deep-layer total nitrogen content among different sites were small, all at relatively low levels. The available phosphorus content ranged from 0.1 to 26.8 mg/kg, with an average value of 6.27 mg/kg. At most sites, the available phosphorus content was low in the surface soil but showed high values in the deep soil, which may be related to the promotion of phosphorus activation by the deep distribution of Pinus koraiensis roots. The readily available potassium content ranged from 65 to 500 mg/kg, with an average value of 140 mg/kg. In summary, the soil in the case area has good nutrient retention capacity, creating a superior nutrient environment for the deep root development of Pinus koraiensis, effectively promoting its growth and development, and providing an important guarantee for the high quality and high yield of Korean pine seeds.

Tab.3 Statistical table of soil fertility indicator test results

Number

Soil layer depth (cm)

pH

Organic matter (g/kg)

CEC / (cmol(+)/kg)

Total nitrogen (g/kg)

Available phosphorus (mg/kg)

Available zinc (mg/kg)

P1

0-20

5.2

44.6

24.49

0.174

1.6

177

 

20-40

5.0

24.9

25.60

0.113

4.6

109

 

40-60

5.0

8.20

20.40

0.053

9.4

80

 

60-80

5.3

5.55

23.92

0.047

25.8

99

 

80-100

5.3

5.77

25.22

0.053

21.6

109

P2

0-20

5.6

89.3

12.35

0.498

2.7

129

 

20-40

5.5

39.8

11.52

0.241

2.0

105

 

40-60

5.6

18.7

10.66

0.121

3.8

75

 

60-80

5.7

9.69

12.03

0.078

3.0

65

 

80-100

5.3

11.2

17.50

0.056

7.4

75

P3

0-20

5.7

48.8

12.13

0.236

2.5

310

 

20-40

5.7

32.5

14.10

0.207

2.5

249

 

40-60

5.8

20.0

12.07

0.127

3.6

181

 

60-80

5.7

17.2

13.69

0.115

4.9

225

 

80-100

5.8

14.8

14.10

0.094

12.0

156

P4

0-20

5.2

29.4

15.38

0.133

2.5

136

 

20-40

5.5

10.9

10.33

0.087

1.6

90

 

40-60

5.4

6.72

12.49

0.051

2.9

87

 

60-80

5.5

14.1

16.25

0.061

4.5

96

 

80-100

5.6

9.10

16.15

0.055

6.8

92

P5

0-20

6.3

43.4

17.38

0.253

1.9

500

 

20-40

6.5

34.6

13.53

0.192

1.6

440

 

40-60

6.7

26.8

13.14

0.145

4.5

251

 

60-80

6.8

12.0

10.76

0.073

8.2

100

 

80-100

6.4

5.45

8.98

0.051

3.9

96

P6

0-20

5.1

30.7

18.84

0.190

0.1

66

 

20-40

5.4

27.9

20.90

0.187

0.6

81

 

40-60

5.3

17.6

19.72

0.107

0.4

68

 

60-80

5.3

5.29

22.14

0.060

4.4

101

 

80-100

5.3

5.45

27.24

0.058

10.2

133

P7

0-20

5.5

35.6

19.10

0.191

0.7

86

 

20-40

5.7

21.2

14.57

0.168

1.4

77

 

40-60

5.7

21.0

15.33

0.149

1.8

79

 

60-80

5.8

10.5

9.07

0.080

2.6

66

 

80-100

5.8

5.31

10.65

0.049

7.8

78

P8

0-20

5.4

63.2

26.20

0.381

2.1

160

 

20-40

5.3

37.8

22.23

0.234

2.2

128

 

40-60

5.4

18.6

12.21

0.136

4.5

91

 

60-80

5.7

17.1

13.49

0.112

11.4

96

 

80-100

5.8

10.2

14.06

0.077

18.0

92

P9

0-20

5.4

70.0

31.84

0.479

7.3

146

 

20-40

5.4

53.6

26.38

0.370

8.0

129

 

40-60

5.5

42.7

22.90

0.270

10.5

121

 

60-80

5.6

19.5

21.13

0.128

15.6

164

 

80-100

5.8

7.54

23.09

0.050

26.8

310

 

111土壤肥力图7

Fig.6 Soil organic matter, total nitrogen, available phosphorus, and readily available potassium contents in different soil layers

(2) Soil environmental quality status

The soil environmental indicator test results were evaluated according to the national standard "Soil Environmental Quality Risk Control Standard for Soil Contamination of Agricultural Land (Trial)" (GB15618-2018)[11]. The soil environmental quality of the Pinus koraiensis forest in Lushuihe Town is excellent. All tested indicators were below the national soil pollution risk screening values, and no pesticide residues were detected, fully meeting the soil environmental quality requirements for the production area of Pinus koraiensis growth[11]. The excellent soil environmental quality provides a safety guarantee for the healthy growth of Pinus koraiensis and helps to produce high-quality and safe forest products.

Tab.4 Statistical table of soil environmental indicator test results

Pollutant Item

Risk Screening Limit

( GB 15618—2018)

Unit

Testing result

P1

P2

P3

P4

P5

P6

P7

P8

P9

Plumbum (Pb)

90

mg/kg

26

26

25

22

19

23

24

25

23

Cadmium (Cd)

0.3

mg/kg

0.20

0.28

0.25

0.22

0.22

0.14

0.18

0.19

0.20

Chromium (Cr)

150

mg/kg

75

67

73

92

87

80

61

75

69

Arsenic (As)

140

mg/kg

10.1

8.99

9.08

8.77

8.20

8.97

8.56

8.2

8.22

Hydrargyrum (Hg)

1.8

mg/kg

0.064

0.070

0.059

0.052

0.052

0.056

0.054

0.058

0.063

Cuprum (Cu)

50

mg/kg

25

19

19

26

28

15

15

17

20

Zinc (Zn)

200

mg/kg

89

97

112

73

75

82

85

97

100

Nickel (Ni)

70

mg/kg

27

27

32

32

35

26

22

25

28

HCHs (Total)

ND

ug/kg

ND

ND

ND

ND

ND

ND

ND

ND

ND

DDTs (Total)

ND

ug/kg

ND

ND

ND

ND

ND

ND

ND

ND

ND

 

3.2.5 Vegetation Cover and Land Use Status

The Normalized Difference Vegetation Index (NDVI) is an important indicator for characterizing surface vegetation growth status. For this dataset, Sentinel data with a 10 m resolution for 2025 were selected as the data source to enhance the spectral response of vegetation and reduce interference from non-vegetation factors, generating the NDVI analysis map of Lushuihe Town (Fig.7). The NDVI in Lushuihe Town generally presents a spatial distribution pattern dominated by high values, with most values concentrated above 0.6, indicating high regional vegetation coverage and good ecological conditions, exhibiting typical forest landscape characteristics of the Changbai Mountain forest region. From a spatial structure perspective, the NDVI shows a mosaic pattern combining continuously distributed high-value backgrounds with scattered low-value patches. The low-value areas are mainly distributed in punctate or clumped forms, mostly corresponding to areas with concentrated human activities such as residential areas, forest logging areas, and open valley areas. Meanwhile, NDVI exhibits strong positive spatial autocorrelation, with high-value areas showing contiguous distribution and weak overall heterogeneity, though local areas exhibit significant differences due to human disturbance.

In the land use classification study, the authors selected Sentinel remote sensing images with a 10 m resolution and a cloud cover percentage of less than 1% in 2025 as the data source. Through steps including image correction, sample extraction for each land use type, supervised classification, and stratified sampling accuracy testing, the land use status of Lushuihe Town was analyzed and constructed (Fig.8). The land use types in Lushuihe Town mainly include construction land, forest land, cultivated land, water bodies, etc. Among them, forest land accounts for approximately 82.54%, cultivated land accounts for approximately 10.04%, and construction land accounts for approximately 5.04%. Forest shade regulates the microclimate, fertile volcanic soil supplies nutrients, and clean water sources maintain physiological metabolism, jointly shaping the characteristics of Korean pine seeds, including their large and plump size, rich oil content, and excellent flavor.

 

 

 

 

 

 

 

 

 

 

Fig.7 NDVI analysis map of Lushuihe Town

Fig.8 Land use distribution map of Lushuihe Town

3.2.6 Distribution Status of Pinus koraiensis Forest

Forest lands in Lushuihe Town generally have Pinus koraiensis distribution, involving forest farms such as Dongsheng, Hongguang, Xinxing, Yongqing, Xilin, and Liming. With reference to the protection scope map of the Lushuihe Korean Pine Seed Kernels Geographical Indication Product, the forest zoning map of Lushuihe Forestry Bureau of Jilin Province (1:100,000), and the key areas for pine seed harvesting, the authors delineated the main concentrated distribution areas of Pinus koraiensis with high density (Fig.9). The main concentrated distribution areas of Pinus koraiensis in Lushuihe are located in the surrounding areas of Lushuihe Town center, Hongguang in the west, and Dongsheng Forest Farm in the south, covering an area of nearly 20,000 hm².

 

 

 

 

 

 

 

 

 

Fig.9 Main concentrated distribution areas of Pinus koraiensis in Lushuihe Town

3.3 Product Characteristic Data

3.3.1 Characteristics of Fusong Lushuihe Korean Pine Seeds

The seeds of Fusong Lushuihe Korean Pine Seeds require low-temperature stratification for more than 120 days to germinate, with a natural germination rate of less than 5%. Pinus koraiensis needs to grow for more than 50 years before it can produce seeds, and the seeds mature only once every two years, resulting in a scarce yield. The seed kernels are light yellow, oval-shaped, plump and uniform, with a thin light brown skin on the surface. The kernel flesh is milky white, fine and oily in texture. They have good storage durability and can maintain their flavor and quality for a long time under low-temperature and dry environments. They are highly resistant to oxidation and not prone to rancidity, possessing good nutritional and health values as well as strong adaptability for food processing.

3.3.2 Quality Test and Analysis of Fusong Lushuihe Korean Pine Seeds

Regarding the quality of Fusong Lushuihe Korean Pine Seeds, this study collected pine seed samples harvested in 2025 and manually opened, mainly sourced from pine seeds from areas around the town center and Dongsheng Forest Farm. The samples were entrusted to the Jilin Provincial Quality Supervision and Inspection Institute and Qingdao Huace Testing Technology Co., Ltd. for testing and analysis. Referring to the "Product of Geographical IndicationLushuihe Korean Pine Seed Kernels" (GB/T 195052008) [14], the testing was conducted in three aspects: sensory indicators, physicochemical indicators, and safety and hygiene indicators [14]. Among them, sensory indicators included four testing items: color, odor, appearance, and impurities. Physicochemical indicators included 16 testing items such as fat, acid value, peroxide value, protein, amino acids, vitamins, and minerals. Safety and hygiene indicators included nine contaminant content items such as arsenic, lead, mercury, cadmium, aflatoxin, and coliform bacteria.

(1) Sensory Indicators

Fusong Lushuihe Korean Pine Seed kernels have a rich and fragrant pine aroma. The kernels are yellow in color, with uniform color and no obvious color difference. The grains are plump and uniform in size. The proportion of slightly damaged kernels is common, while severely damaged kernels account for less than 3%, and broken kernels account for less than 7%. The proportions of severely damaged and broken kernels are kept at relatively low levels, reflecting that effective measures have been taken during the harvesting, processing, storage, and transportation of pine seeds to ensure the overall appearance quality of the product. The texture is fine, with no visible impurities. During processing, procedures such as impurity screening and cleaning have been properly implemented, resulting in a clean and hygienic product (Tab.5), which meets the requirements of the national standard [14].

Tab.5 Statistical table of sensory indicator test results of Fusong Lushuihe Korean Pine Seeds

Number

Pollutant item

Standard Indicator

Test results

Judgment

1

Colour and lustre

Yellow

Yellow

Compliant

2

Smell

Strong pine scent

Strong pine scent

Compliant

3

Appearance

Minor damage < 3%, broken kernels < 7%

Minor damage < 3%, broken kernels < 7%

Compliant

4

 Impurities

No visible impurities

No visible impurities

Compliant

(2) Physicochemical Indicators

Korean pine seeds are recognized as a high-nutrition nut. Due to its unique growing environment, Fusong Lushuihe Korean Pine Seeds have even more prominent nutritional components. The test results (Tab.6) show that the crude fat content is 65.4%, and the dietary fiber content is 8.4 mg/100g. The high fiber content complements its high fat content beneficially. The acid value is 0.6 mg/g, and the peroxide value is 0.0023 g/100g. The extremely low acid value and peroxide value indicate a low degree of oxidative rancidity, resulting in better taste and nutritional value. The protein and amino acid contents are abundant, reaching 16.7 g/100g and 13.1 g/100g, respectively, making them a high-quality plant protein source. The amino acids include 16 types: aspartic acid, threonine, serine, glutamic acid, glycine, alanine, valine, methionine, isoleucine, leucine, tyrosine, phenylalanine, lysine, histidine, arginine, and proline. The vitamin E content is relatively high, at 24.7 g/100g. Among the major elements, the calcium content is 145 mg/100g, potassium content is 594 mg/100g, and magnesium content is 2.58×10³mg/100g. Among the trace elements, the manganese content is 56.1 mg/100g, iron content is 45.5 mg/100g, and zinc content is 62.7 mg/100g.

Tab.6 Statistical table of physicochemical indicator test results of Fusong Lushuihe Korean Pine Seeds

Number

Pollutant item

Unit

Standard Indicator

Test results

Judgment

1

Fat

%

60

65.4

Compliant

3

Acid Value

mg/g

4

0.60

Compliant

4

Peroxide Value

g/100g

0.08

0.0023

Compliant

5

Protein

g/100g

ND

16.7

ND

6

Amino Acids

g/100g

ND

13.1

ND

7

 Vitamin E

g/100g

ND

24.7

ND

8

Vitamin B₁

g/100g

ND

0.40

ND

9

Vitamin B₂

g/100g

ND

0.0551

ND

10

Calcium

mg/100g

ND

145

ND

11

Potassium

mg/100g

ND

594

ND

12

 Magnesium

mg/100g

ND

2.58×103

ND

13

Manganese

mg/100g

ND

56.1

ND

14

Iron

mg/100g

ND

45.5

ND

15

Zinc

mg/100g

ND

62.7

ND

16

 Dietary Fiber

mg/100g

ND

8.4

ND

 

(3) Safety and Hygiene Indicators

The test results of Fusong Lushuihe Korean Pine Seeds show that the contents of heavy metals such as arsenic, lead, cadmium, and total mercury were either not detected or far below the national standard limits. Aflatoxin Bwas not detected, and the counts of coliform bacteria and other microorganisms were low, demonstrating excellent microbial safety performance (Tab.7). All test results comply with the standards of the "National Food Safety StandardLimits of Contaminants in Food" (GB 27622022)[12] and the "National Food Safety StandardLimits of Mycotoxins in Food" (GB 27612017) [13]. This indicates that the safety and hygiene indicators of Fusong Lushuihe Korean Pine Seeds fully meet the national standards, and that their growth and processing processes strictly adhere to safety specifications, ensuring the safety of the product.

Tab.7 Statistical table of hygiene indicator test results of Fusong Lushuihe Korean Pine Seeds

Number

Pollutant item

Unit

Standard Indicator

Test results

Judgment

1

 Arsenic

mg/kg

0.5

ND

Compliant

2

Lead

mg/kg

0.5

ND

Compliant

3

Cadmium

mg/kg

0.5

0.103

Compliant

4

 Mercury

mg/kg

ND

ND

ND

5

Aflatoxin B1

μg/kg

10

ND

Compliant

6

Coliform Bacteria

CFU/g

ND

10

ND

7

 Mould and Yeast

CFU/g

ND

7.8×104

ND

8

Salmonella

/25g

ND

ND

ND

9

Staphylococcus aureus

CFU/g

ND

10

ND

 

4 Social and Economic Development, Business Management, and Historical Traditions of Lushuihe Town , Fusong County

4.1 Population and socio-economic development situation

The county economy of Fusong County has achieved steady and rapid progress, and this sound economic condition has laid the foundation for the comprehensive advancement of agricultural green development throughout Fusong County. In 2025, the gross domestic product (GDP) of Fusong County exceeded 15 billion yuan, and the total output value of agriculture, forestry, animal husbandry, and fishery in the county increased by 5.0% year-on-year. The per capita disposable income of rural residents in Fusong County reached 22,008 yuan, surpassing the average level of Jilin Province. After the national complete cessation of commercial logging, Lushuihe Town has undergone a profound industrial transformation, shifting from a single focus on forest logging and wood processing to a diversified industrial structure encompassing natural mineral water, ginseng, and pine nut kernels, while actively developing ecotourism based on national forest parks and hunting grounds, committing to a green and sustainable development path[15–18]. Relying on over one million centuries-old mother forests of Pinus koraiensis and a well-established “planting–harvesting–processing–sales” industrial chain, the pine nut kernel industry of Lushuihe Town has become the leading “one town, one specialty” industry of Fusong County, with annual sales exceeding 200 million yuan and exports accounting for over 60%. This industry directly employs more than 3,000 local forest farmers, achieving a three-dimensional synergistic development of “ecological resource protection, traditional skill inheritance, and industrial economic growth.” In 2025, the output of pine nut kernels in Lushuihe Town reached approximately 4,500 tons, generating over 200 permanent jobs and more than 400 seasonal jobs, with a monthly wage of 4,500 yuan.

4.2 Protection and Development of Pinus koraiensis Resources

Since the 1980s, with the increasing awareness of natural forest protection, the Lushuihe Forestry Bureau took the lead in establishing a research group dedicated to the conservation and utilization of Pinus koraiensis resources. This group systematically advanced research on the cultivation of mother forests, improvement of cone yield, and optimization of kernel quality. Between 1990 and 1995, utilizing a technical system comprising “plus tree selection – artificial pollination – seedling domestication,” the research team successfully screened three superior mother tree strains of Pinus koraiensis. Subsequently, a large-scale mother forest renovation project was carried out in the core forest area of Lushuihe, expanding the protected area for ancient trees over one hundred years old to 120,000 mu (8,000 hm²)[19]. In 2003, “Lushuihe Pinus koraiensis Kernels” were officially approved as a National Geographical Indication Product, becoming the first nut product in Northeast China to receive this certification. Over the following two decades, through continuous scientific research efforts, the products successively obtained Grade AA Green Food Certification, EU Organic Food Certification (EU Organic), and China Forest Certification (CFCC)[20]. In 2020, Lushuihe Pinus koraiensis mother forest kernels were included in the second batch of the China-EU Geographical Indications Agreement list. In 2021, they were recognized as “China Famous Agricultural Products,” and the traditional technique of collecting Pinus koraiensis nuts was inscribed in the Jilin Provincial Intangible Cultural Heritage List. In 2023, the Lushuihe Ancient Pinus koraiensis Forest in Jilin Province was successfully selected as one of “China’s Most Beautiful Ancient Tree Groves.”

4.3 Fusong Lushuihe Pine Nut Management

4.3.1 Pine Nut Resource Control

Relying on Asia's largest natural mother forest of Pinus koraiensis, Lushuihe Town implements refined management of Pinus koraiensis resources. Based on the growth cycle and seed-bearing regularity of Pinus koraiensis, a strict quota-based collection system is enforced, with the annual collection volume determined according to monitoring data to ensure the sustainable growth of mother trees. Forestry departments and forest farms have established a comprehensive monitoring system for Pinus koraiensis resources, integrating satellite remote sensing, unmanned aerial vehicle (UAV) monitoring, and ground patrols to obtain real-time information on the growth status, pest and disease occurrence, and forest ecological environment changes of Pinus koraiensis.

4.3.2 Integrated management model combining industry, academia, research, government, cooperatives, local residents, and traditional practices

Lushuihe Town has developed an integrated management model that combines industry, academia, research, government, cooperatives, local residents, and traditional practices. The government provides active guidance to foster collaboration between research institutions and enterprises, facilitating the selection and breeding of improved Pinus koraiensis varieties, optimization of shell-breaking and kernel-extraction processes, and research on the functional components of pine nuts. Enterprises have actively introduced internationally advanced aseptic production lines and processing technologies. In the primary processing stage, impurities and defective kernels are precisely removed, and low-temperature roasting is employed to maximize the retention of nutritional components and flavor of the pine nut kernels. Cooperatives are responsible for organizing standardized harvesting, drying, and preliminary sorting by villagers, ensuring the purity and traceability of the raw materials. By participating in harvesting, management, and transportation, villagers not only increase their income but also provide a stable supply of high-quality raw materials to enterprises.

Leveraging the geographical indication advantages of Lushuihe pine nuts as a product of origin, the government has actively promoted the clustered development of the industry and the extension of the industrial chain through policy guidance and investment attraction. This has gradually led to the formation of a modern industrial system encompassing multiple links, including wild resource management, integrated mechanical harvesting, low-temperature shell breaking and kernel extraction, deep processing of oils, and development of health foods. Meanwhile, with leading enterprises such as Jilin Changbai Mingzhu Forest Food Co., Ltd. playing a key role, multi-party efforts have been made to strengthen the research and development of "pine nut +" series products, resulting in composite products combining pine nuts with ginseng, cocoa, crispy candies, and other ingredients, thereby continuously extending the industrial chain.

4.3.3 Market Operation Strategies

The government fully plays a leading role, guiding enterprises through industrial policies to precisely align with consumer demand and develop diversified products targeting different consumer groups. For the high-end consumer market, organically certified pine nut products with exquisite gift-box packaging are introduced, emphasizing ecological, healthy, and high-quality attributes. For ordinary consumers, affordable, small-packaged, ready-to-eat pine nut kernels are offered to meet daily consumption needs. Meanwhile, the government actively attracts investment and introduces professional institutions for e-commerce operations and brand planning, helping enterprises utilize emerging marketing tools such as social media and live streaming to narrate the brand story and ecological value of Lushuihe pine nuts, thereby enhancing consumer awareness and purchase intention.

In terms of brand building, the government leads the formulation of regional public brand standards for "Lushuihe Pinus koraiensis Pine Nuts" and collaborates with industry associations to strengthen brand authorization and supervision. Brand promotion emphasizes the natural ecological characteristics and high nutritional value. Through integrated online and offline communication, exhibition participation, promotional events, and cultural tourism integration projects, the brand visibility and market share are continuously enhanced. Concurrently, market regulatory authorities, in conjunction with industry associations, strengthen the protection of the geographical indication brand "Lushuihe Pinus koraiensis Pine Nut Kernels," regularly conduct special rectification campaigns, and rigorously combat counterfeit and substandard products to safeguard brand reputation.

4.4 Inheritance and Promotion of Fusong Lushuihe Pine Nut Culture

The history of Pinus koraiensis in Lushuihe Town can be traced back to ancient times. Numerous forestry studies indicate that Pinus koraiensis, as an ancient tree species, has existed in East Asia for at least ten thousand years, with a distribution range even broader in the past than at present. Local ethnic minorities in Fusong County, such as the Manchu and Korean ethnic groups, have coexisted with Pinus koraiensis since ancient times[21]. During the Qing Dynasty, Manchu aristocrats regarded the pine nuts of Pinus koraiensis as a delicacy. Not only were they consumed as everyday food, but they also played a crucial role in important ceremonies such as sacrificial rites. Pine nut kernels were listed as one of the "Eight Delicacies" and were often used in the preparation of pastries and other offerings. Among the common people, pine nut kernels have been an integral part of traditional dietary culture, being made into specialty foods such as pine nut porridge and pine nut cakes, passed down through generations. Pinus koraiensis holds a special place in local culture, symbolizing longevity and tenacity. In folk legends, it is regarded as the embodiment of the mountain deity, guarding the forests and the people living there.

Diverse cultural activities centered on Pinus koraiensis have been developed. The government plays a guiding and supportive role, coordinating multiple departments including forestry, cultural tourism, and market regulation to ensure comprehensive management throughout event planning, safety assurance, and brand promotion, thereby creating the Pinus koraiensis Cultural Festival. During the festival, traditional processing techniques of pine nuts are demonstrated, specialty foods are tasted, and folk performances and knowledge contests related to Pinus koraiensis are held. These activities help preserve and promote Pinus koraiensis culture, enhance local residents' awareness of protecting Pinus koraiensis resources and their cultural identity, and enable more people to appreciate the unique regional cultural charm of Lushuihe pine nuts[21]. Furthermore, the integrated development of "Pinus koraiensis culture + eco-tourism" is actively promoted. Leveraging Lushuihe's locational advantages adjacent to Fusong County town, Erdaobaihe Town, and the core scenic area of Changbai Mountain, outdoor activities such as forest hunting cultural experiences, wilderness hiking, mountain camping, and forest exploration are combined. High-quality tourism routes are jointly launched with surrounding scenic spots."

4.5 Traceability of Pine Nut Habitat in Lushuihe, Fusong

Promoting the sustainable development of the pine nut industry in Lushuihe, Fusong, requires robust support from modern technology. To this end, a landmark habitat ground station has been established in the forest farm of the case study area (Figure 10) to enable real-time, round‑the‑clock monitoring and traceability of the ecological conditions of the Pinus koraiensis forest. The key indicators recorded include negative oxygen ions, phenology, wind speed, wind direction, rainfall, air quality, temperature, relative humidity, soil temperature and moisture, atmospheric pressure, and soil electrical conductivity. These real‑time monitoring data can reflect the high‑quality growing environment of the pine nuts and provide scientifically accurate data support for phenological changes and related responses of the Pinus koraiensis forest. Consequently, this facilitates a more intelligent, scientific, and sustainable transformation of the cultivation model of the pine nut industry.


Fig.10 Diagram of the Lushuihe Town Pine Nut Geotope Habitat Ground Station Facilities

5 Conclusions

The Fusong Lushuihe Pine Nut is produced in the core area of Changbai Mountains, Fusong County, Jilin Province. The pristine forest ecosystem, characterized by high altitude, cold temperate humid climate, and fertile volcanic ash soil, provides favorable habitat conditions for the growth of Pinus koraiensis. The soil environmental quality in Lushuihe Town is excellent, with no detectable pesticide residues, and the clean water quality facilitates the production of high-quality pine nuts. Lushuihe Pine Nuts exhibit unique qualities including plump kernels, milky white nut meats, and rich aroma, and contain substantial amounts of fat, protein, and essential trace elements for the human body, demonstrating significant nutritional value. Local authorities have achieved protective development of the precious Pinus koraiensis mother forest resources by regulating the collection and management of wild pine nuts, and by constructing and developing an integrated management model incorporating industry, academia, government, community, and traditional practices. Future efforts should further deepen the integration of ecological protection and industrial development, enhancing the quality and added value of pine nuts while ensuring the security and stability of Pinus koraiensis resources, thereby achieving synergistic improvements in ecological, economic, and social benefits. The Case Study on Habitat Protection and Sustainable Development of Pinus koraiensis Forests for Fusong Lushuihe Pine Nuts aims to provide a scientific pathway for the conservation of rare tree species resources and high-quality industrial development, contributing to the practical deepening of the concept that “lucid waters and lush mountains are invaluable assets.”

 

Author contributions: Qi, W. made the overall design of this case, including the overall design of the dataset, the construction of the content framework of the case study paper, the technical route, and the writing design, and collaboratively convened case field investigations and seminars, and carried out paper writing and coordination. Wang, G., Gao, J. T., Du, Y. C., and Zhao, W. X. coordinated the case investigation and data collection. Qi, W., Zhu, W. H., Zhu, J. H., Chen, Y. T., Liu, Y., Liu, Y. B., and Niu, Z. M. participated in the field investigation of the case study. Wang, X. Q., Jin, X., and Qi, W. were responsible for the collection and analysis of soil and water samples. Chen, Y. T., Liu, Y., Liu, Y. B., Niu, Z. M., Li, Y., and Fan, K. were responsible for the collection, analysis, and processing of data and the production of figures. Tang, X. R. and Liu, H. Q. were responsible for coordinating assistance from units at various levels and the collection of local cultural data. Qi, W. was responsible for the collection and analysis of meteorological data. Zhu, J. H. was responsible for the collection and analysis of culture related to Korean pine seeds. Liu, H. Q., Xu, Y. G. and Zhang, Z. G. were responsible for the collection of management, economic development, and characteristic data. Zhu, W. H., Zhu, J. H., Chen, Y. T., Liu, Y., Liu, Y. B., Niu, Z. M., Li, Y., Fan, K., and Tang, X. R. participated in the writing of the paper.

Acknowledgements: We would like to express our gratitude to Professor Liu, C., Song, X. F., and Wang, Z. B. of the Institute of Geographic Sciences and Natural Resources Research, Chinese Academy of Sciences for their guidance and assistance in the completion of this case data and paper. We also thank the villagers who provided help during the collection and analysis of soil and water samples.

Conflict of Interest Statement:

The authors declare no conflicts of interest


References:

[1] Liu, J. B. (2012). Management and protection measures of natural Pinus koraiensis mother forest in Lushuihe. Jilin Forestry Science and Technology, 41(3), 44-45.

[2] Zu, S. C. (2020). Resource evaluation and deep processing technology of Pinus koraiensis Sieb. et Zucc. seeds [Master's thesis]. Harbin: Northeast Forestry University.

[3] Li, X. H. (2019). Study on the relationship between vegetation productivity and water use in the broad-leaved Pinus koraiensis forest region of Changbai Mountain [Doctoral dissertation]. Beijing: Beijing Forestry University.

[4] Liu, D. D. (2017). Research progress on selective breeding of Pinus koraiensis in China. Protection Forest Science and Technology, (3), 96-99+116.

[5] Zhang, H. N., Guo, J. Q., & Zhao, Y. H. (2022). Research status on composition, nutritional value, development and utilization of Pinus koraiensis seeds. Forest By-Product and Specialty in China, (2), 84-86.

[6] Song, W., Yan, P. Y., Liu, J., et al. (2024). Case study on habitat protection and sustainable development of Qinxian Millet in Shangdang Basin. Journal of Global Change Data, 8(3), 267-285+386-404.

[7] Global Change Research Data Publishing System. (2017). Global change research data sharing policy [OL]. https://doi.org/10.3974/dp.policy.2014.05 (Updated 2017).

[8] Qi, W., Zhu, J. H., Wang, G., et al. (2026). Fusong Lushuihe Korean Pine Seeds Pinus koraiensis Forest Geographical Indication Habitat Case Dataset [J/DB/OL]. Digital Journal of Global Change Data Repository, 2026. https://doi.org/10.3974/geodb.2026.04.01.V1.

[9] Li, H. Y., Sun, Y. H., Lu, L. L., et al. (2013). Analysis of water resources characteristics in Fusong Drinking Water Source Protection Area. Jilin Geology, 32(3), 80-85.

[10] Ministry of Ecology and Environment of the People's Republic of China, & State Administration for Market Regulation. (2021). Standard for irrigation water quality (GB 5084—2021) [S]. Beijing: Standards Press of China.

[11] Ministry of Ecology and Environment of the People's Republic of China, & State Administration for Market Regulation. (2018). Soil environmental quality—Risk control standard for soil contamination of agricultural land (Trial) (GB 15618—2018) [S]. Beijing: Standards Press of China.

[12] National Health Commission of the People's Republic of China, & State Administration for Market Regulation. (2022). National food safety standard—Limits of contaminants in food (GB 2762—2022) [S]. Beijing: Standards Press of China.

[13] National Health and Family Planning Commission of the People's Republic of China, & China Food and Drug Administration. (2017). National food safety standard—Limits of mycotoxins in food (GB 2761—2017) [S]. Beijing: Standards Press of China.

[14] General Administration of Quality Supervision, Inspection and Quarantine of the People's Republic of China, & Standardization Administration of China. (2008). Product of geographical indication—Lushuihe Korean pine seed kernels (GB/T 19505—2008) [S]. Beijing: Standards Press of China.

[15] Wang, X. H. (2019). Discussion on the construction technology of Pinus koraiensis fruit timber forest. Agriculture and Technology, 39(3), 88-89.

[16] Wang, W., Li, P. X., Wang, G. X., et al. (2009). Effects of storage temperature on the physiology and quality of pine nut kernels under different states. Zhejiang Agricultural Science, (4), 743-746.

[17] Dong, S. N., Yu, Y. F., & Fu, Y. C. (2023). Study on the spatiotemporal differentiation and influencing factors of the quality of under-forest economic growth in Changbai Mountain area of Jilin Province. Journal of Northeast Normal University (Philosophy and Social Sciences Edition), (1), 147-158+202.

[18]  Liang, Y. F. (2024). Evaluation of development level and model effect analysis of agricultural green development pilot zone in Fusong County [Master's thesis]. Changchun: Jilin University.

[19]  Ren, Z. J., & Shi, W. H. (2024). Exploration of forest resource protection and development under the background of sustainable development—A case study of Fusong County. Guangdong Sericulture, 58(10), 24-26.

[20] Liu, L., & Nie, H. Y. (2018). Analysis of the protection development status and countermeasures of national geographical indication products in Jilin Province. Science and Technology Innovation Herald, 15(28), 152-153.

[21]  Xu, W. (2019). "Shanlihong" forever red—The growth track of Shanlihong Troupe in Fusong County. Shenhua (Part 2), (7), 62-65.



[1] SRTM data source: https://www.earthdata.nasa.gov/data/instruments/srtm